Research Interests

• Origin of oxygenic photosynthesis
• Light harvesting systems in oxygenic algae and plants
• Carotenoid biosynthesis and protection
• Red algal development (Nori relatives)

Awards

• Fellow of the American Association for the Advancement of Science, 1982 
• National Academy of Science, 1996 
• Distinguished University Professor at the University of Maryland, 1997 
• Fellow of American Society of Plant Biologists, 2007 
• Phycological Society of America Award of Excellence, 2011

Education

• Ph.D., Northwestern University, 1963
• B.A., Blackburn College, 1958